The usual model construction for NFU preserves information

The usual construction of models of NFU (New Foundations with urelements, introduced by Jensen) is due to Maurice Boffa. A Boffa model is obtained from a model of (a fragment of) Zermelo-Fraenkel with Choice (ZFC) with an automorphism which moves a rank: the domain of the Boffa model is a rank that is moved. "Most" elements of the domain of the Boffa model are urelements in terms of the interpreted NFU. The main result of this paper is that the restriction of the membership relation of the original model of set theory with automorphism to the domain of the Boffa model is first-order definable in the language of NFU. In particular, all information about the extensions in the original model of the urelements of the model of NFU is definable in terms of NFU. A corollary (answering a question of Thomas Forster) is that the urelements in a Boffa model are not homogeneous.